Abundance: 0.02 to 47.47% (median 0.91%)
Latitudinal range: -28.2° to 44.4°Habitats: Mediterranean woodland (3), desert/xeric shrubland (1), temperate broadleaf/mixed forest (10), temperate savanna (5), tropical/subtropical dry broadleaf forest (12), tropical/subtropical moist broadleaf forest (35), tropical/subtropical savanna (9)
Found in 75 samples
Cameroon: La Belgique (sleeping sites)
Ecuador: Centro Cientifico de Rio Palenque (May 1975), Centro Cientifico de Rio Palenque (February 1976), Centro Cientifico de Rio Palenque (July 1976)
Gabon: Ipassa Reserve (large rainy season)
Ghana: Ankasa Conservation Area (unlogged forest), Ankasa Conservation Area (plantation)
India: Thirunelly Forests, Kurukshetra, Singhori Wildlife Sanctuary (mixed forests site 2)
Indonesia: Napu Valley (natural forest), Palolo Valley (annual cultures), Toro (natural forest/anoa dung), Napu Valley (young secondary forest), Napu Valley (agroforestry ecosystem), Napu Valley (annual culture), Palolo Valley (natural forest), Toro (monospecific planted shade trees/anoa dung)
Italy: Chiusa Pesio
Ivory Coast: Lola Field Station (forest)
Japan: Northeastern Tochigi Prefecture, Shiraoi (cattle dung), Mukawa (cattle dung)
Malaysia: Temengor Forest Reserve (logger camp), Danum Valley Field Centre (Riverine A), Sungkai Reserve Forest (secondary forest)
Mexico: Boca del Chajul, Los Tuxtlas (coati dung/nocturnal), Los Tuxtlas (continuous forest), Los Tuxtlas (howlers present), Los Tuxtlas (howlers absent), Santa Rosa, San Martín Volcano (forest fragments), Vicente Guerrero (IF), El Jaralillo Cattle Ranch (July), San Martín Volcano (forest-pasture edges), San Martín Volcano (pastures), Los Tuxtlas (mosaic habitat), Los Tuxtlas (forest fragments), San Fernando (tropical deciduous forest), San Fernando (pasture with Acacia shrubs)
Mozambique: Maputo Special Reserve (reserve grassland), Maputo Special Reserve (reserve farm)
Panama: El Hato del Volcan (small traps)
Portugal: Odivelas
South Africa: Hluhluwe Game Reserve, Skukuza (control)
Spain: Salamanca (spring)
Tanzania: Majawanga (bush site 1), Majawanga (field site 4)
Thailand: Ton Nga Chang Wildlife Sanctuary (primary forest), Ton Nga Chang Wildlife Sanctuary (secondary forest), Mo Singto Long Term Ecological Research plot (macaque), Mo Singto Long Term Ecological Research plot (barking deer), Mo Singto Long Term Ecological Research plot (sambar deer), Mo Singto Long Term Ecological Research plot (elephant)
Tunisia: Mejerda Valley
Turkey: Dagmarmara (600 m, 2004)
Uganda: Kibale National Park (Ngogo), Kibale National Park (Pinus plantation)
United States: Blackbeard Island, Cumberland Island, East of Arapaho Prairie, Arapaho Prairie, Welder Wildlife Refuge (open chaparral), Hutcheson Memorial Forest disturbed field (June/July), Hutcheson Memorial Forest old growth forest (June/July), Lake City Research Center Site A, Lake City Research Center Site B, Brookings County (cattle manure), Highland Road Observatory Park (March), Missouri Ozark Forest Ecosystem Project (no harvest management)
Vietnam: Ba Be National Park (dung beetles)
See also Copris acutidens, Copris agnus, Copris amyntor, Copris camerunus, Copris cariniceps, Copris carinicus, Copris carmelita, Copris davisoni, Copris denticulatus, Copris doriae, Copris elphenor, Copris fricator, Copris hispanus, Copris imitans, Copris inhalatus, Copris jahi, Copris laeviceps, Copris lugubris, Copris lunaris, Copris macacus, Copris mesacanthus, Copris minutus, Copris moechus, Copris orphanus, Copris pecuarius, Copris phungae, Copris puncticollis, Copris punctulatus, Copris ramosciceps, Copris ramosiceps, Copris reflexus, Copris repertus, Copris sallei, Copris saundersi, Copris serius, Copris signatus, Copris sinicus, Copris sp. 4, Copris sp. 6, Copris sp. 7, Copris sp. B, Copris spinator, Copris truncatus, Copris tullius, Copris typhoeus